Those values are not passing a data validation or data type, or maybe even an edit_If constraint (Which to be honest I don’t think it is the case because the whole row wouldn’t be recorded at all instead of it recording partially as you describe.)
possibility 2. If the error is persisting just with that particular record then it is indeed not finding any record for those last values. (Which I think is the most likely thing happening)
Edit: Just to be sure, try regenerating both tables, sometimes the error is in there and don’t know why but recent updates won’t mark this type of errors.
Remove all occurrences of [_THISROW]. in your expression–none of them are needed. I have experienced times where its inclusion produced unexpected results.